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Dave Foley Flik (voice)
Kevin Spacey Hopper (voice)
Julia Louis-Dreyfus Atta (voice)
Hayden Panettiere Dot (voice)
Phyllis Diller Queen (voice)
Richard Kind Molt (voice)
David Hyde Pierce Slim (voice)
Joe Ranft Heimlich (voice)
Denis Leary Francis (voice)
Jonathan Harris Manny (voice)
Madeline Kahn Gypsy Moth (voice)
Bonnie Hunt Rosie (voice)
Michael McShane Tuck / Roll (voice)
John Ratzenberger P.T. Flea (voice)
Brad Garrett Dim (voice)
Roddy McDowall Mr. Soil (voice)
Edie McClurg Dr. Flora (voice)
Alex Rocco Thorny (voice)
David Ossman Loco (voice)
Jack Angel Thud (voice)
Bob Bergen Aphie / Male Ants (voice)
Kimberly J. Brown Female Ants (voice) (as Kimberly Brown)
Rodger Bumpass Harry Mosquito (voice)
Anthony Burch Female Ants (voice)
Rachel Davey Baby Maggots (voice)
Paul Eiding Male Ants (voice)
Jessica Evans Female Ants (voice)
Bill Farmer Male Ants (voice)
Sam Gifaldi Ants (voice)
Brad Hall Grasshopper (voice)
Jess Harnell Bus Bettle (voice)
Brenden Hickey Thumper (voice) (as David Lander)
John Lasseter Bug Zapper Bug #1 (voice)
Sherry Lynn Female Ants (voice)
Mickie McGowan Fly (voice)
Christina Milian Female Ants (voice)
Kelsey Mulrooney Female Ants (voice)
Ryan O'Donohue Grasshopper (voice)
Phil Proctor Fly Boy (voice)
Jan Rabson Axel (voice)
Jordan Ranft Female Ants (voice) (as Franchesca Smith)
Andrew Stanton Bug Zapper Bug #2 (voice)
Hannah Swanson Female Ants (voice)
Russi Taylor Female Ants (voice) (as Russie Taylor)
Travis Tedford Ant Kids (voice)
Ashley Tisdale Lead Blueberry Scout (voice)
Lee Unkrich Ant #1 (voice)
Jordan Warkol Ant #2 (voice)
India De Maio Ants (uncredited)
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